Maximum Yield USA 2012 March | Page 134

Six Ways Plants Move Tropisms are plant movements directed toward or away from a stimulus." developmental response to water) and thermotropism (response dependent upon temperature). It is helpful to also designate plant responses that are triggered by a definite stimulus, but not in a direction that is related to the direction of the stimulus. Such movements are called nastic movements. Examples of this are a mimosa folding its leaves in response to a touch or a Dionaea (Venus flytrap) closing in response to the presence of an insect. A lot of tropism research has focused on how plant hormones known as auxins control plant growth by stimulating cell elongation. It is well accepted that phototropic and geotropic bending of shoots and roots results from cells on one side of a plant elongating faster than cells on the other side, thus causing the plant to bend and change the direction of its growth. 132 Maximum Yield USA | March 2012 Phototropism Photosynthesis is a popular way to make a living on Earth and plants—as well as some microscopic organisms—have finely tuned, light-controlled positioning systems. Auxins are important in regulating how plant organs move and grow toward a light source and photosynthesis is one of the most-studied topics in botany. In the basic model of phototropism, the concentration of auxin is elevated on the unlit side of a plant shoot exposed to light, which causes the cell walls on that side to become less fixed in structure. The cells on the unlit side elongate and then re-solidify their cell walls—the effect of this deceptively simpleseeming process is that the elongating cells cause the shoot to bend toward the light. Exactly how the concentration of auxin gets to be higher on the dark side is not perfectly understood, but current research suggests that auxin is transported from one side to the other and perhaps from other places in the plant. The biochemistry of auxin-regulated bending is still being worked out, even after many decades of research. Two other pieces of the phototropism puzzle lie in the proteins phototropin and phytochrome. Although these two molecules are almost certainly on the light-receiving end of the phototropism system, the biochemical steps that go from there to the action of auxin are still under investigation.